作者单位海南师范学院生物系, 海口 571158; 中国热带农业科学院热带作物生物技术国家重点实验室, 海口 571101[1]海南师范学院生物系, 海口 571158[2]西南师范大学生命科学学院, 重庆 400715[3]中国热带农业科学院热带作物生物技术国家重点实验室, 海口 571101[4]Department of Biology,Hainan Normal College,Haikou 571158,China;National Key Biotechnology Laboratory for Tropical Crops,Chinese Academy of Tropical Agricultural Sciences,Haikou 571101,China[5]Department of Biology,Hainan Normal College,Haikou 571158,China[6]College of Life Science,Southwest China Normal University,Chongqing 400715,China[7]National Key Biotechnology Laboratory for Tropical Crops,Chinese Academy of Tropical Agricultural Sciences,Haikou 571101,China[8]