作者单位:Graduate School of Bioagricultural Sciences, Nagoya University, Chikusa, Nagoya, Aichi 464-8601, Japan[1]Graduate School of Biosphere Science, Hiroshima University, Higashi-Hiroshima, Hiroshima 739-8528, Japan[2]Japanese Avian Bioresource Project Research Center, Hiroshima University, Hiroshima 739-8528, Japan[3]