作者单位:Univ Calif San Francisco, Neurosci Grad Program, San Francisco, CA 94143 USA[1]Univ Calif San Francisco, Weill Inst Neurosci, San Francisco, CA 94143 USA[2]Univ Calif San Francisco, Dept Psychiat & Behav Sci, San Francisco, CA 94143 USA[3]Univ Calif San Francisco, Sch Med, San Francisco, CA USA[4]