作者单位:South African National Biodiversity Institute, Private Bag X7, Claremont, Cape Town 7735, South Africa[1]Department of Botany and Zoology, DST-NRF Centre of Excellence for Invasion Biology, University of Stellenbosch, Private Bag X1, Matieland 7602, South Africa[2]Natural History Division, Iziko South African Museum, P.O. Box 61, Cape Town 8000, South Africa[3]