作者单位:Centre for Ecology and Conservation, University of Exeter, Penryn, Cornwall, TRIO 9EZ, UK,ProDelphinus, Lima, Peru[1]Centre for Ecology and Conservation, University of Exeter, Penryn, Cornwall, TRIO 9EZ, UK[2]Centre for Ecology and Conservation, University of Exeter, Penryn, Cornwall, TRIO 9EZ, UK,Environment and Sustainability Institute, University of Exeter, Penryn,Cornwall, UK[3]