作者单位:Scripps Institution of Oceanography and Department of Neurosciences, University of California, San[1]Department of Biological Sciences, Vanderbilt University, 465 21st Ave S, Nashville, TN 37240[2]Mote Marine Laboratory, 1600 Ken Thompson Parkway, Sarasota, FL 34236, United States[3]Department of Physiology, Development and Neuroscience, University of Cambridge, Anatomy Building[4]