作者单位:Nippon Paper Ind Co Ltd, Agribiotechnol Res Lab, Kita Ku, 5-21-1 Oji, Tokyo 1140002, Japan[1]Univ Tsukuba, Tsukuba Plant Innovat Res Ctr, Tsukuba, Ibaraki 3058752, Japan[2]Univ Tsukuba, Grad Sch Life & Environm Sci, Tsukuba, Ibaraki 3058752, Japan[3]