作者单位:Shimane University, Department of Biological Science, Faculty of Life and Environmental Science, 1060 Nisikawatsu-cho, Matsue, Shimane 690-8504, Japan[1]Tottori University, Division of Resources Life Science, The United Graduate School of Agricultural Sciences, 4-101 Koyama-Minami, Tottori 680-8553, Japan[2]