作者单位:Department of Biophysics, University of MichiganAnn Arbor, MI, United States[1]Department of Biophysics, University of MichiganAnn Arbor, MI, United States, Department of Cell[2]Department of Biomedical Engineering, University of MichiganAnn Arbor, MI, United States[3]Department of Cell and Developmental Biology, University of MichiganAnn Arbor, MI, United States[4]