作者单位:Neurobiology, Neurodegeneration and Repair Laboratory, National Eye Institute, Bethesda, MD 20892, United States[1]Department of Vision Science, University of Alabama, Birmingham, Birmingham, AL 35924, United States[2]Verna and Marrs McLean Department of Biochemistry and Molecular Biology, Baylor College of Medicine, Houston, TX 77030, United States[3]