作者单位:Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48104, USA[1]School of Natural Resources and the Environment, University of Michigan, Ann Arbor, MI 48104, USA[2]Institute of Integrative Biology, ETH, Zurich 8007, Switzerland[3]Department of Entomology, University of California, Davis, CA 95616, USA[4]