作者单位：Department of Plant and Wildlife Biology, Brigham Young University, Provo, UT 84602, USA[1]Rancho Santa Ana Botanic Garden, 1500 N College Avenue, Claremont, CA 91711, USA[2]US Forest Service, Inyo National Forest, 351 Pacu Ln, Bishop, CA 93514, USA[3]White Mountain Research Station, 3000 E Line St, Bishop, CA 93514, USA[4]Natural Science Division, Pepperdine University, Malibu, CA 90263, USA[5]
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