作者单位:Laboratoire d’Adaptation des Plantes aux Stress Abiotiques, Centre de Biotechnologie, Technopole de Borj-Cedria (CBBC), B.P. 901, 2050 Hammam-Lif, Tunisia[1]Dipartimento di Produzione Vegetale, University of Milan, Via Celoria 2, I-20133 Milano, Italy[2]