作者单位：Department of Biological Applications and Technology, Laboratory of Zoology, University of Ioannina, University Campus of Ioannina, Ioannina, 45110, Greece[1]Laboratoire Ecologie des Systemes Marins Cotiers UMR 5119, Universite Montpellier 2, cc 093, Place Eugene Bataillon, Montpellier Cedex 5, 34095, France[2]
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