作者单位:Institute of Lowland and Marine Research, Saga University, Saga, Saga, 840-8512, Japan[1]Nishi-Abashiri Fishermen’s Association, Abashiri, Hokkaido, 093-0045, Japan[2]Department of Aquatic Bioscience, Tokyo University of Agriculture, Abashiri, Hokkaido, 099-2493, Japan[3]Laboratory of Benthology, Graduate School of Fisheries Sciences, Hokkaido University, Hakodate, Hokkaido, 041-8611, Japan[4]