作者单位:Institute of Botany, Academy of Sciences of the Czech Republic, CZ-25243 Pruhonice 1,Czech Republic[1]Institute of Botany, Academy of Sciences of the Czech Republic, CZ-25243 Pruhonice 1, Czech Republic,and Herbarium Universitatis Carolinae Pragensis, Benátská 2, CZ-12801 Praha, Czech Republic[2]