作者单位:Univ Sao Paulo, Fac Ciencias Farmaceut, Dept Ciencias Farmaceut, Ribeirao Preto, Brazil[1]Univ Sao Paulo, Fac Filosofia Ciencias & Letras, Dept Biol, BR-14040901 Ribeirao Preto, SP, Brazil[2]Univ Sao Paulo, Inst Biociencias, Dept Ecol, Sao Paulo, Brazil[3]