作者单位:Wildlife Conservation Society, Amenabar 1595, C1426AKC Ciudad de Buenos Aires, Argentina[1]Biological Sciences, Institute of Environmental Sustainability, University of Wales, Swansea, SA2 8PP, UK[2]Centro Nacional Patagónico (CENPAT), CONICET, Boulevard Brown 2915, U9120ACD Puerto Madryn, Chubut, Argentina[3]