作者单位:Gyeongsang Natl Univ, Div Appl Life Sci Program BK21Plus, Plant Mol Biol & Biotechnol Res Ctr, Jinju 52828, South Korea[1]Vietnam Natl Univ Agr, Fac Biotechnol, Hanoi, Vietnam[2]Dong Eui Univ, Dept Life Sci & Biotechnol, Busan 47340, South Korea[3]