作者单位:Botany Section, Department of Environmental Biology, University of Siena, Via P.A. Mattioli 4, 53100 Siena, Italy[1]Laboratoire d'Adaptation des Plantes aux Stress Abiotiques, Centre de Biotechnologie, Technopole de Borj Cédria, BP 901, Hammam-Lif 2050, Tunisia[2]