作者单位:Universite Montpellier 2-CNRS, Institut des Sciences de l'Evolution, 00065, 34095 Montpellier cedex 05, France[1]Department of Botany and Zoology, Faculty of Science, Masaryk University, Koticii'skd 2, 611 37 Brno, Czech Republic[2]Laboratoire de Mathematique Physique et Systemes (EA 4217), Universite de Perpignan, Via Domitia,[3]