作者单位：California Department of Fish and Game, 407 West Line Street, Bishop, CA 93514, USA[1]Department of Fish, Wildlife, and Conservation Biology, Colorado State University, Fort Collins, CO 80523, USA[2]Colorado Division of Wildlife, 317 West Prospect Road, Fort Collins, CO 80526, USA[3]Colorado Division of Wildlife, 2300 South Townsend Avenue, Montrose, CO 81401, USA[4]
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