作者单位:Guangdong Entomological Institute, 105 Xingang Road W., 510260 Guangzhou, China[1]Graduate University of Chinese Academy of Sciences, Beijing 100039, China[2]College of Life Sciences, Sun Yat-sen University, 510275 Guangzhou, China[3]South China Botanical Garden of Chinese Academy of Sciences, Guangzhou 510650, China[4]