作者单位：Graduate School of Fisheries Science and Environmental Studies, Nagasaki University, Bunkyo 1-14, Nagasaki 852-8521,Japan[1]Department of Biological Sciences, College of Science, Sungkyunkwan University, Suwon 440-746, South Korea[2]Department of Marine Science, College of Natural Sciences, Incheon National University, Incheon 406-772, South Korea[3]
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