作者单位Research Unit Palaeontology, Ghent University, Krijgslaan 281/S 8, 9000 Ghent, Belgium;Research Assistant of the Fund for Scientific Research-Flanders(F. W. O.-Vlaanderen)[1]中国科学院南京地质古生物研究所,现代古生物学和地层学国家重点实验室 南京 210008[2]Research Unit Palaeontology, Ghent University, Krijgslaan 281/S 8, 9000 Ghent, Belgium[3]