摘要As important subunits of the leading-strand DNA polymerase epsilon,chromatin remodeling,and histone acetylation complexes,the H2A/H2B-like histone-fold domain-containing proteins DNA PO-LYMERASE II SUBUNIT B3(DPB3)and DPB4 play key roles in nucle-osome assembly and heterochromatin maintenance during DNA replication in yeast,Drosophila,and mammals(He et al.,2017;Bellelli et al.,2018;Yu et al.,2018;Casari et al.,2021).DPB3 and DPB4 were usually classified as NUCLEAR FACTOR Y(NF-Y)family proteins in plants,but less is known for their functions(Petroni et al.,2012;Laloum et al.,2013).In Arabidopsis,DPB3-1/NF-YC10 inter-acts with the transcription factor DEHYDRATION-RESPONSIVE ELEMENT BINDING PROTEIN2A(DREB2A)to enhance heat stress-inducible gene expression via a transcriptional complex with NF-Y subunits NF-YA2 and NF-YB3(Sato et al.,2014).OsDPB3-2 in rice(Oryza sativa)functions similarly to AtDPB3-1(Sato et al.,2016).A recent study reported that AtDPB3-1 interacts with glyceral-dehyde-3-phosphate dehydrogenase and regulates transcriptional and physiological responses to heat stress(Kim et al.,2020).