江苏高校优势学科建设工程资助项目, 农业部农作物保护项目江苏省自然科学基金面上项目江苏省高校自然科学研究重大项目(12KJA180001)资助. This study was supported by the Project Funded by the Priority Academic Program Development of Jiangsu Higher Education Institutions, the Project on Crop Conservation Funded by the Ministry of Agriculture of Chinathe General Projects Funded by Natural Sci-ence Foundation of Jiangsu Provincethe Major Project Funded by Natural Science Foundation of Jiangsu Higher Education Institutions