作者单位Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering,National Observations and Research Station for Wetland Ecosystems of the Yangtze Estuary,Institute of Biodiversity Science,School of Life Sciences,Fudan University,Shanghai 200438,China[1]Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering,National Observations and Research Station for Wetland Ecosystems of the Yangtze Estuary,Institute of Biodiversity Science,School of Life Sciences,Fudan University,Shanghai 200438,China;Research Center for Ecology,College of Science,Tibet University,Lhasa 850000,China;Tibet University—Fudan University Joint Laboratory for Biodiversity and Global Change,Fudan University,Shanghai 200032,China[2]Ecology,Department of Biology,University of Konstanz,78464 Konstanz,Germany[3]