摘要The pol en tube is fundamental for the reproduction of seed plants. Characteristical y, it grows relatively quickly and uni‐directional y (“polarized growth”) to extend the male gametophyte to reach the female gametophyte. The pol en tube forms a channel through which the sperm cel s move so that they can reach their targets in the ovule. To grow quickly and directional y, the pol en tube requires an intense movement of organel es and vesicles that al ows the cel ’s contents to be distributed to sustain the growth rate. While the various organel es distribute more or less uniformly within the pol en tube, Golgi‐released secretory vesicles accumulate massively at the pol en tube apex, that is, the growing region. This intense movement of organel es and vesicles is dependent on the dynamics of the cytoskeleton, which reorganizes differential y in response to external signals and coordinates membrane trafficking with the growth rate of pol en tubes.